Ikora offered 2-3 Story Missions, rotating on a weekly basis. Choosing a mission locked the player out of the others until the one they picked was completed. Interacting with a device next to Ikora started the mission. Upon completion, the player received an Upgrade Point, if they still had some skills to unlock, and 3 Vanguard Research Tokens.
Meditations were replaced with a new mode in Forsaken. The Daily Heroic Story Missions mode from the Vanguard Director served the same purpose, but with no reward, until it was removed in Beyond Light.